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''Homo antecessor'' is an archaic human species of the Lower Paleolithic, known to have been present in Spain, and possibly England and France, between about 1.2 million and 0.8 million years ago (Mya). It was described in 1997 by Eudald Carbonell, Juan Luis Arsuaga and José María Bermúdez de Castro, who based on its "unique mix of modern and primitive traits" classified it as a previously unknown archaic human species. The fossils associated with ''Homo antecessor'' represent the oldest direct fossil record of the presence of ''Homo'' in Europe. The species name ''antecessor'' proposed in 1997 is a Latin word meaning "predecessor", or "vanguard, scout, pioneer". Authors who do not accept ''H. antecessor'' as a separate species consider the fossils in question an early form of ''H. heidelbergensis'' or as a European variety of ''H. erectus''.

Taxonomy



Research history

''H. antecessor'' was discovered in 1991 in the Gran Dolina site of the Sierra de Atapuerca region of northern Spain by Spanish palaeoanthropologists José María Bermúdez de Castro, Eudald Carbonell, and Juan Luis Arsuaga. These remains were deposited in the sixth level (TD6) dating to about 780,000 years ago, making them the oldest European human fossils. Stone tools indicate hominin activity in Europe as early as 1.6 mya in Eastern Europe and Spain. The sediment of Gran Dolina was dated at 900,000 years old in 2014. More than 80 bone fragments from six individuals were uncovered in 1994 and 1995. The site also had included approximately 200 stone tools and 300 animal bones. Stone tools including a stone carved knife were found along with the ancient hominin remains. All these remains were dated at least 900,000 years old. The best preserved specimens are ATD6-15 and ATD6-69, a frontal bone and a maxilla (upper jawbone) of a 10 year old boy, dubbed the "Gran Dolina Boy" (''el chico de la Gran Dolina''), dating to 859–782,000 years ago. Direct ESR dating of ATD6-92 in 2018 resulted in an estimated age of 949 to 624 thousand years ago, further restricted by magnetostratigraphic data from TD6 unit of Atapuerca Gran Dolina to 949 to 772 thousand years ago. In 2007, a molar dating to 1.2–1.1 million years ago was recovered from the nearby Sima del Elefante site. It belonged to a 20–25 year old individual. In 2008, an additional mandible fragment, stone flakes, and evidence of butchery were discovered. Until 2013 with the discovery of the 1.4 Ma infant tooth from Barranco León, Orce, Spain, these were the oldest human fossils known from Europe. Evidence of early human presence in England and France has later been tentatively associated with ''H. antecessor'' purely on chronological grounds and not based on anatomical evidence. Fifty footprints dating to between 1.2 million and 800,000 years ago were discovered in Happisburgh, England. They were possibly made by an ''H. antecessor'' group.

Classification

''H. antecessor'' has been proposed as a chronospecies intermediate between ''H. erectus'' (c. 1.9–1.4 Mya) and ''H. heidelbergensis'' (c. 0.8–0.3 Mya). While ''H. heidelbergensis'' is widely accepted as the immediate predecessor of ''H. neanderthalensis'', and possibly ''H. sapiens'', the derivation of ''H. heidelbergensis'' from ''H. antecessor'' is debatable. ''H. antecessor's'' discoverers suggested ''H. antecessor'' as a derivation of African ''H. erectus'' (''H. ergaster'') which would have migrated to the Iberian Peninsula at some point before 1.2 Mya, and developed into ''H. heidelbergensis'' by 0.8 Mya, and further into ''H. neanderthalensis'' after 0.3 Mya. In 2009, palaeoanthropologist Richard Klein stated he was skeptical that ''H. antecessor'' was ancestral to ''H. heidelbergensis'', interpreting ''H. antecessor'' as a "failed attempt to colonize southern Europe". The legitimacy of ''H. antecessor'' as a separate species has also been questioned because the fossil record is fragmentary, and especially as no complete skull has been found, with only fourteen fragments and lower jaw bones known. Because of this, it is also proposed that ''H. antecessor'' was an early form of ''H. heidelbergensis'', which would extend the range of ''H. heidelbergensis'' to 1.2–0.3 Mya. A 2020 analysis by Welker et al. of ancient proteins collected from a tooth of an ''H. antecessor'' specimen indicated that ''H. antecessor'' belonged to a "sister lineage" related to the ancestor of modern humans, Neanderthals, and Denisovans, but was not itself their ancestor, corroborating earlier opinions that ''H. heidelbergensis'' was not derived from ''H. antecessor''.

Physiology

Based on the radial and clavicle lengths, the statures of 2 specimens were calculated to be and . The body proportions of ''H. antecessor'' fall within the range of variation for modern humans, and upper limb proportions were likely more similar to those of modern humans than Neanderthals. The facial anatomy of the Boy of Gran Dolina appears very similar to that of modern humans, and it was thus suggested that this marked the beginning of modern human facial features. These features likely did not change so drastically with age. However, modern humanlike facial features may have evolved multiple times independently among different human lineages. Based on teeth eruption pattern, the researchers think that ''H. antecessor'' had the same development stages as ''H. sapiens'', though probably at a faster pace. Other significant features demonstrated by the species are a protruding occipital bun, a low forehead, and a lack of a strong chin. Some of the remains are almost indistinguishable from the fossil attributable to the 1.5-million-year-old Turkana Boy, belonging to ''H. ergaster''. A study on the scapula of 850,000-year-old remains of a ''H. antecessor'' child and an adolescent, which were unearthed at the Grand Dolina site, concluded that they were not skilled climbers, yet they were physically able to throw stones or spears accurately.

Behaviour

The large animal carcasses at Gran Dolina appear to have been carried to the site intact. This indicates that ''H. antecessor'' groups dispatched multi-member hunting parties who delayed eating immediately after a successful kill in order to share it with all group members after hauling it back, showing social cooperation, division of labour, and food sharing. A total of 16 species were recorded from Gran Dolina, including the bush-antlered deer, an extinct species of fallow deer, an extinct red deer, an extinct bison, a wild boar, the rhino ''Stephanorhinus etruscus'', the Stenon zebra, a mammoth, the Mosbach wolf, the fox ''Vulpes praeglacialis'', the Gran Dolina bear, the spotted hyena, and a lynx. The most common butchered remains are those of deer. Also, adult and child ''H. antecessor'' specimens from Gran Dolina exhibit cut marks, crushing, burning, and other trauma indicative of cannibalism, and are the second-most common remains bearing evidence of butchering. It is unclear if cannibalism was practiced often (ritual cannibalism) or if this was an isolated incident in a dire situation (survival cannibalism).

See also

* Ceprano Man * Happisburgh footprints * ''Homo heidelbergensis'' * Out of Africa I * Peopling of Europe * Tautavel Man * Human evolution * Homo

References



External links


Homo antecessor



Early Human Phylogeny

Spanish National Research Centre for Human Evolution (CENIEH)

Atapuerca

Human Timeline (Interactive)
Smithsonian, National Museum of Natural History (August 2016). {{DEFAULTSORT:Homo Antecessor Category:1994 archaeological discoveries Category:Early species of Homo Category:Lower Paleolithic Category:Pleistocene primates Category:Prehistoric Spain Category:Prehistoric cannibalism Category:Fossil taxa described in 1997